[Systematics and Taxonomy]


Benthic Polychaetes of British Columbia and Washington

by J.M. (Lobo) Orensanz

Introduction

[Apologies for the lack of 'pretty' formatting. This is a temporary version - G. Read]



Family APHRODITIDAE Malmgren, 1867
		Genus APHRODITA Linnaeus, 1758
			Aphrodita japonica Marenzeller, 1879
			Aphrodita longipalpa Essenberg, 1917
			Aphrodita negligens Moore, 1905
			Aphrodita parva Moore, 1905
		Genus LAETMONICE Kinberg, 1855
			Laetmonice pellucida Moore, 1903

Family POLYNOIDAE Kinberg, 1856
	Subfamily ARCTONOINAE Hanley, 1989
		Genus ARCTNONOE Chamberlin, 1920
			Arctonoe fragilis (Baird, 1863)
			Arctonoe pulchra (Johnson, 1897)
			Arctonoe vittata (Grube, 1855)
	Subfamily HARMOTHOINAE Horst, 1917
		Genus ARCTEOBIA Annenkova, 1937
				[Genus included in B&H's key]
***			Arcteobia anticostiensis (McIntosh, 1874)
				Fournier & Levings, 1982: Syllogeus 40: 18
***			Arcteobia spinielytris Annenkova, 1937
				Fournier & Levings, 1982: Syllogeus 40: 18-19
		Genus BYLGIDES Chamberlin, 1919
				Pettibone, 1993: Smith. Contr. Zool. 545
*			Bylgides macrolepidus (Moore, 1905)
				Pettibone, 1993: Smith. Contr. Zool. 545: 21-23, f.13
				[= Antinoella m. in B&H]
		Genus EUNOE Malmgren, 1865
			Eunoe depressa Moore, 1905
			Eunoe nodosa (Sars, 1861)
			Eunoe oerstedi Malmgren, 1865
			Eunoe senta (Moore, 1902)
			Eunoe uniseriata Banse & Hobson, 1968
		Genus GATTYANA McIntosh, 1900
			Gattyana ciliata Moore, 1902
			Gattyana cirrosa (Pallas, 1766)
			Gattyana treadwelli Pettibone, 1949
		Genus GAUDICHAUDIUS Pettibone, 1986
				Pettibone, 1986: Smith. Contr. Zool. 428: 33-40
*			Gaudichaudius iphionelloides (Johnson, 1901)
				[= Gattyana i. in B&H]
				Pettibone, 1986: Smith. Contr. Zool. 428: 37-40, f.18
		Genus HARMOTHOE Kinberg, 1855
			Harmothoe extenuata (Grube, 1840)
			Harmothoe fragilis Moore, 1910
			Harmothoe imbricata (Linnaeus, 1767)
			Harmothoe multisetosa (Moore, 1902)
		Genus HERMADION Kinberg, 1855
			Hermadion truncata (Moore, 1902)
		Genus HESPERONOE Chamberlin, 1919
***			Hesperonoe adventor (Skogsberg, 1928)
				[Included in B&H's keys]
				Fournier & Levings, 1982: Syllogeus 40: 19-20
			Hesperonoe complanata (Johnson, 1901)
		Genus MALMGRENIELLA Hartman, 1967
				Pettibone, 1993: Smith. Contr. Zool. 538.
***			Malmgreniella bansei Pettibone, 1993
				Pettibone, 1993. Smith. Contr. Zool. 538: 63-65, f.42
***			Malmgreniella berkeleyorum Pettibone, 1993
				Pettibone, 1993. Smith. Contr. Zool. 538: 63, f.41
				[= Harmothoe lunulata in B&B 1945, 1948]
***			Malmgreniella liei Pettibone, 1993
				Pettibone, 1993. Smith. Contr. Zool. 538: 65-68, f.43
**			Malmgreniella nigralba (E.Berkeley, 1923)
				Pettibone, 1993. Smith. Contr. Zool. 538: 60-63, fs.39-40
				[= Harmothoe lunulata in B&H] 
***			Malmgreniella scriptoria Pettibone, 1993
				Pettibone, 1993. Smith. Contr. Zool. 538: 68, f.44
		Genus TENONIA Nichols, 1969
**			Tenonia priops (Hartman, 1961)
				Barreca, 1984: Proc. Biol. Soc. Wash. 97: 801-803
				[= T. kitsapensis in B&H]
	Subfamily LEDIDASTHENIINAE Pettibone, 1989
		Genus LEPIDASTHENIA Malmgren, 1867
			Lepidasthenia berkeleyae Pettibone, 1948
			Lepidasthenia longicirrata Berkeley, 1923
	Subfamily LEPIDONOTINAE Willey, 1902
		Genus HALOSYDNA Kinberg, 1855
			Halosydna brevisetosa Kinberg, 1855
		Genus LEPIDONOTUS Leach, 1816
			Lepidonotus squamatus (Linnaeus, 1767)
	Subfamily POLYNOINAE Kinberg, 1856
		Genus POLYEUNOA McIntosh, 1885
			Polyeunoa tuta (Grube, 1855)
		Genus POLYNOE Savigny, 1818
			Polynoe canadensis (McIntosh, 1874)
			Polynoe gracilis (Verrill, 1874)
	Unnamed Subfamily
		Genus HOLOLEPIDA Moore, 1905
			Hololepida magna Moore, 1905

Family PHOLOIDAE Kinberg, 1858
				Pettibone, 1992: Smith. Contr. Zool. # 532, 24 p.
		Genus PHOLOE Johnston, 1839
				Pettibone, 1992: Smith. Contr. Zool. # 532: 4.
Q			Pholoe sp. 
				As "Pholoe minuta" in B&H. Records of that species correspond to a variety of other Pholoids; citations from the Pacific NW need validation.
		Genus PHOLOIDES Pruvot, 1895
				Pettibone, 1992: Smith. Contr. Zool. # 532: 15-16
*			Pholoides asperus (Johnson, 1897) 
				[as Peisidice a. in B&H] 
				Pettibone, 1992: Smith. Contr. Zool. # 532: 18-20

Family SIGALIONIDAE Malmgren, 1867
		Genus SIGALION Audouin & Milne-Edwards (in Cuvier, 1830)
				[Included in B&H's keys]
				Mackie & Chambers, 1990: Zool. Scr. 19: 39-56
***			Sigalion mathildae Audouin & Milne-Edwards (in Cuvier, 1830)
				Fournier & Levings, 1982: Syllogeus 40: 20
				Mackie & Chambers, 1990: Zool. Scr. 19: 40-45, fs.1-5
		Genus STHENELAIS Kinberg, 1855
			Sthenelais berkeleyi Pettibone
			Sthenelais tertiaglabra Moore, 1910

Family CHRYSOPETALIDAE Ehlers, 1864
		Genus DYSPONETUS Levinsen, 1879
				[Included in B&H's keys]
***			Dysponetus pygmaeus Levinsen, 1879
				This species has never been formally cited for the Pacific NW, but was included in Kozloff (1987). It is common in the San Juan Islands, among laminarian holdfasts.
		Genus PALEANOTUS Schmarda, 1861
			Paleanotus bellis (Johnson, 1897)

Family AMPHINOMIDAE Lamarck, 1818
		Genus CHLOEIA Lamarck, 1818
			Chloeia entypa Chamberlin, 1919

Family EUPHROSINIDAE Williams, 1851
				Kudenov, 1993: Antarct. Res. Ser. 58: 93-150
		Genus EUPHROSINE Lamarck, 1818
				Kudenov, 1987: Bull. Biol. Soc. Wash. 7: 184-193.
				Kudenov, 1993: Antarct. Res. Ser. 58: 104-107.
			Euphrosine arctia Johnson, 1897
			Euphrosine bicirrata Moore, 1905
			Euphrosine heterobranchia Johnson, 1901
			Euphrosine hortensis Moore, 1905

Fmily LACYDONIIDAE Bergstrom, 1914
***			Unidentified species
				Fournier & Levings, 1982: Syllogeus 40: 21.

Family PHYLLODOCIDAE Oersted, 1843
				Pleijel, 1991: Zool. Scr. 20: 225-261. Definition of subfamilies proposed by Pleijel (based on a cladistic analysis, followed below) differs radically from the traditional system.
				Pleijel, 1993: POLYCHAETA PHYLLODOCIDAE. Marine Invertebrates of Scandinavia, No. 8, 158 p.
	Subfamily NOTOPHYLLINAE Pleijel, 1991
		Genus BERGSTROEMIA Banse, 1973
				Status under review by Eibye-Jacobsen; may be a synonym of Austrophyllum Bergstrom (Eibye-Jacobsen, 1991: 130; Pleijel, 1991: 253; op.cit.).
Q			Bergstroemia nigrimaculata (Moore, 1909)
				[= Eulalia (B.) n. in B&H]
				Eibye-Jacobsen, 1991: Steenstrupia 17: 130
		Genus NOTOPHYLLUM Oersted, 1843
				Pleijel, 1991: Zool. Scr. 20: 233-234, f.4
			Notophyllum imbricatum Moore, 1906
		Genus NEREIPHYLLA
				Pleijel, 1991: Zool. Scr. 20: 235, fs.7, 22C-F
*			Nereiphylla castanea (Marenzeller, 1879)
				[= Phyllodoce (Genetyllis) c. in B&H]
	Subfamily PHYLLODOCINAE Oersted, 1843
				Pleijel, 1991: Zool. Scr. 20: 235.
		Genus PHYLLODOCE Lamarck, 1818
				Pleijel, 1988: Zool. Scr. 17: 141-153
				Pleijel, 1991: Zool. Scr. 20: 238, f.10.
				Anonymous, 1995: Pigmentation of Phyllodoce maculata, mucosa and williamsi. Pacific NW Polychaete Workshop, informal document, one page.
			Phyllodoce citrina Malmgren, 1865
				Pleijel, 1988: Zool. Scr. 17: 143, f.2
				Pleijel, 1993: Marine Invert. Scandinavia 8: 35-37, fs.20-21
			Phyllodoce groenlandica Oersted, 1842
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 14-15
				Pleijel, 1988: Zool. Scr. 17: 144-145, f.3.
				Pleijel, 1993: Marine Invert. Scandinavia 8: 37-40, fs.22-23.
***			Phyllodoce hartmanae Blake & Walton, 1977
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 15
***			Phyllodoce longipes (Kinberg, 1866)
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 15-16
				Pleijel, 1988: Zool. Scr. 17: 146-147, f.5.
			Phyllodoce maculata (Linnaeus, 1767)
				Pleijel, 1988: Zool. Scr. 17: 147-149, f.6.
				Pleijel, 1993: Marine Invert. Scandinavia 8: 48-51, fs.31-32.
**			Phyllodoce medipapillata Moore, 1909
				[= P. madeirensis in B&H]
				McCammon & Montagne, 1979: Zool. J. Linn. Soc.66: 362-363, f.5a-c
				Mountford, 1991: Ophelia Suppl. 5: 166.
Q			Phyllodoce mucosa Oersted, 1843
				Pleijel, 1988: Zool. Scr. 17: 149-151, f.7.
				Pleijel, 1993: Marine Invert. Scandinavia 8: 51-53, fs.33-34.
				Originally described from Denmark; cited for many Arctic-boreal areas, but non-European records require validation. Easily confused with P. maculata.
***			Phyllodoce multipapillata Kravitz & Jones, 1979
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 14, f.4C-D
			[?] Phyllodoce multiseriata Rioja, 1941
			Phyllodoce williamsii (Hartman, 1936)
		Genus PARANAITIS Southern, 1914
				Pleijel, 1991: Zool. Scr. 20: 237-238, fs.9, 23B.
			Paranaitis polynoides (Moore, 1909)
	Subfamily ETEONINAE sensu Pleijel, 1991
				Pleijel, 1991: Zool. Scr. 20: 240.
		Genus HESIONURA Hartmann-Schröder, 1958
				Pleijel, 1991: Zool. Scr. 20: 247, f.18.
			Hesionura coineaui difficilis (Banse, 1963)
		Genus MYSTIDES Theel, 1879
				Pleijel, 1991: Zool. Scr. 20: 247, f.19
Q			Mystides borealis (Theel, 1879)
				Pleijel, 1993: Marine Invert. Scandinavia 8: 127-129, f.89.
				Originally described from Novaya Zemlya; cited for world-wide areas, but records other than the type locality require validation.
	   Genera in the "EULALIA" group (under PHYLLODOCINAE in the traditional system)
		Genus EULALIA Savigny, 1822
				Pleijel, 1991: Zool. Scr. 20: 241-243, fs.15, 22C.
Q			[?] Eulalia bilineata (Johnston, 1840)
				Pleijel, 1993: Marine Invert. Scandinavia 8: 89-91, fs.62-63. Originally described from the UK; cited for world-wide areas.
				Harris, 1995: Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages. Records from the Pacific coast of North America correespond to several other species.
				Harris, 1995: Comparison of European and Eastern Pacific Eulalia bilineata. Pacific NW Polychaete Workshop, informal document, one page.
			Eulalia levicornuta Moore, 1909
			Eulalia parvoseta Banse & Hobson, 1968
				Originally described as E. (Pterocirrus); probably Eulalia s. str. (Pleijel, 1991: Zool. Scr. 2O: 256)
			Eulalia quadrioculata Moore, 1906
Q			Eulalia viridis (Linnaeus, 1767)
				Pleijel, 1993: Marine Invert. Scandinavia 8: 110-113, fs.79-81.
				Originally described from "Oceano septentrionali"; cited for world-wide areas, but records outside the North Atlantic require validation.
		Genus EUMIDA Malmgren, 1865
				Eibye-Jacobsen, 1991: Steenstrupia 17: 81-140
				Pleijel, 1991: Zool. Scr. 20: 241, f.13.
			Eumida longicornuta (Moore, 1906)
				Eibye-Jacobsen, 1991: Steenstrupia 17: 108-110
Q			Eumida sanguinea (Oersted, 1843)
				Eibye-Jacobsen, 1991: Steenstrupia 17: 87-93
				Pleijel, 1993: Marine Invert. Scandinavia 8: 76-78, fs.52-54.
				Originally described from Denmark; cited for world-wide areas, but records outside Europe and Arcto-Atlantic areas require validation.
		Genus PTEROCIRRUS Claparède, 1868
				Pleijel, 1991: Zool. Scr. 20: 240-241, fs.12, 22A-B.
Q			Pterocirrus macroceros (Grube, 1860)
				Pleijel, 1993: Marine Invert. Scandinavia 8: 61-63, fs.40-42.
				Originally described for the Mediterranean; cited for world-wide areas, but non-European records require validation. At least some records may correspond to P. montereyensis: Harris, 1995, Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages
		Genus SIGE Malmgren, 1865
				[Included in B&H's keys]
				Pleijel, 1990: Zool. J. Linn. Soc. 98: 161-184
***			Sige bifoliata Moore, 1909
				Pleijel, 1990: Zool. J. Linn. Soc. 98: 167-169, f.3
				[Cited for British Columbia by Berkeley, 1924, but not included in CPF or B&H]
	   Genera of ETEONINAE sensu strictum (traditional definition)
		Genus ETEONE Savigny, 1822
				Wilson, 1988: Mem. Mus. Victoria 49: 385-431
				Pleijel, 1991: Zool. Scr. 20: 249, f.21.
***			Eteone californica Hartman, 1936
				[Included in B&H's keys]
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 9
				Wilson, 1988: Mem. Mus. Victoria 49: 390-392, fs.1b-e, 14b
				Fournier & Levings, 1982: Syllogeus 40: 20.
***			Eteone columbiensis Kravitz & Jones, 1979
				Kravitz & Jones, 1979: Bull.So.Calif.Acad. Sci. 78: 10-12, f.3D-G
				Wilson, 1988: Mem. Mus. Victoria 49: 392, fs.2a-d, pl.1a
Q			Eteone longa (Fabricius, 1780)
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 9
				Wilson, 1988: Mem. Mus. Victoria 49: 399-400, fs.4e-h, pl.1d
				Pleijel, 1993: Marine Invert. Scandinavia 8: 143-144, fs.100-101.
				Originally described from Greenland. Records of the E. flava/longa-group are confused, as it may include several different Arctic to boreal species. The group is need of extensive review.
			Eteone pacifica Hartman, 1936
				Wilson, 1988: Mem. Mus. Victoria 49: 400
Q			Eteone spetsbergensis Malmgren, 1865
				Wilson, 1988: Mem. Mus. Victoria 49: 403-405, fgs.6d-g, pl.2a
				Status uncertain, requires redescription (Pleijel, 1993: Marine Invert. Scandinavia 8: 150).
				Records from the Pacific NW may be conspecific with Eteone sp. sensu Kravitz & Jones, 1979: (Bull. So. Calif. Acad. Sci. 78: 12, f.4A-B)
***			Eteone spilotus Kravitz & Jones, 1979
				Kravitz & Jones, 1979: Bull.So.Calif. Acad. Sci. 78: 9-10, f.3A-C
				Wilson, 1988: Mem. Mus. Victoria 49: 405-407, fs.7a-d, 14f
			Eteone tuberculata Treadwell, 1922
				Wilson, 1988: Mem. Mus. Victoria 49: 408
		Genus HYPERETEONE Bergstrom, 1914
				Wilson, 1988: Mem. Mus. Victoria 49: 385-431
				(Considered as a synonym of Eteone by Pleijel, 1991, op.cit.)
***			Hypereteone fauchaldi (Kravitz & Jones, 1979)
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 7, f.2A-G (as Eteone f.)
				Wilson, 1988: Mem. Mus. Victoria 49: 412-414, fs.9a-e, pl.2c
		Genus MYSTA Malmgren, 1865
				[Included in B&H's keys, as a subgenus of Eteone]
				Wilson, 1988: Mem. Mus. Victoria 49: 385-431
				(Considered as a synonym of Eteone by Pleijel, 1991, op.cit.)
***			Mysta barbata Malmgren, 1865
				Kravitz & Jones, 1979: Bull. So. Calif. Acad. Sci. 78: 12-14
				Wilson, 1988: Mem. Mus. Victoria 49: 421-422, fs.12a-d, 14l
				Pleijel, 1993: Marine Invert. Scandinavia 8: 134-136, fs.93-94
	Phyllodocids of questionable status
		Genus HESPEROPHYLLUM Chamberlin, 1919
				Nomen dubium fide Pleijel, 1991: Zool.Scr.20: 249, 257
Q			Hesperophyllum tectum Chamberlin, 1919
				Nomen dubium fide Pleijel, 1991: Zool.Scr.20: 249, 257

Family HESIONIDAE Malmgren, 1867
		Genus HETEROPODARKE Hartmann-Schröder, 1862
***			Heteropodarke sp.
				Formally unreported for the area, but I have found it to be common in sandy beaches of Puget Sound. The type species (H. heteromorpha) has been cited from world-wide areas, including California (Dorsey, 1978: Bull. Southern Calif. Acad. Sci. 77: 82-87). Records may correspond to different species (see for example Perkins, 1984: Proc. Biol. Soc. Wash. 97: 565-572)
		Genus KEFERSTEINIA Quatrefages, 1865
				Perkins, 1984: Proc. Biol. Soc. Wash. 97: 574
			Kefersteinia cirrata (Keferstein, 1862)
		Genus MICROPHTHALMUS Mecznikow, 1865
				[Included in B&H's keys]
***			Microphthalmus coustalini Fournier, 1991
				Fournier, 1991: Bull. Mar. Sci. 48: 208-213
***			Microphthalmus hystrix Fournier, 1991
				Fournier, 1991: Bull. Mar. Sci. 48: 208-213
***			Microphthalmus simplicichaetosus Westheide & Purschke, 1992
				Westheide & Purschke, 1992: Proc. Biol. Soc. Wash. 105: 132-135
***			Microphthalmus sp.
				Fournier, 1991: Bull. Mar. Sci. 48: 208-213
		Genus MICROPODARKE Okuda, 1938
			Micropodarke dubia (Hessle, 1925)
		Genus OPHIODROMUS Sars, 1861
			Ophiodromus pugettensis (Johnson, 1901)
		Genus PODARKEOPSIS Laubier, 1961
**			Podarkeopsis glabrus (Hartman, 1961)
				[= Gyptis brevipalpa in B&H]
				Perkins, 1984: Proc. Biol. Soc. Wash. 97: 579

Family PILARGIDAE Saint-Joseph, 1899
				Salazar-Vallejo, 1986: Cah. Biol. Mar. 27: 193-209
				Fitzhugh & Wolf, 1990: Amer. Mus. Novitates 2992, 16 pp.
				Licher & Westheide, 1994: Mém.Mus.natn.Hist.nat.162: 223-235
	Subfamily SIGAMBRINAE Salazar-Vallejo, 1986
		Genus OTOPSIS Ditlevsen, 1917
			Otopsis longipes Ditlevsen, 1917
		Genus PILARGIS Saint-Joseph, 1899
			Pilargis berkeleyae Monro, 1933
		Genus SIGAMBRA F.Müller, 1858
			Sigambra tentaculata (Treadwell, 1941)

Family SYLLIDAE Grube, 1850
				Garwood, 1991: Ophelia 5 Suppl.: 81-87
	Subfamily EXOGONINAE Rioja, 1925
		Genus BRANIA Quatrefages, 1865
			Brania brevipharyngea Banse, 
		Genus EXOGONE Oersted, 1845
				San Martin, 1991: Bull. Mar. Sci. 49: 715-740
			Exogone (E.) lourei Berkeley & Berkeley, 1938
				Perkins, 1981: Proc. Biol. Soc. Wash. 93: 1092-1094
			Exogone (E.) verugera (Claparède, 1868)
*			Exogone (Parexogone) molesta Banse, 1972
***			Exogone sp. A (n. sp., Kudenov & Harris, in press)
				Exogone sp. in Banse, 1972
				Harris, 1995: Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages
**			Exogone sp. B (n. sp., Kudenov & Harris, in press)
				[= E. gemmifera in B&H]
				Harris, 1995: Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages
		Genus SPHAEROSYLLIS Claparède, 1863
				Riser, 1991: Ophelia Suppl. 5: 209-217.
**			Sphaerosyllis californiensis Hartman, 1966
				Riser, 1991: Ophelia Suppl. 5: 216
Q			Sphaerosyllis hystrix Claparède, 1863
				Originally described from European waters, has been recorded from world-wide areas. Non-European records need revision, and most likely correspond to other species.
**			Sphaerosyllis sp. F
				[as S. brandhorsti in B&H]
				Harris, 1995: Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages
**			Sphaerosyllis sp. P, Harris, unpubl.
				[= S. pirifera in B&H]
				Harris, 1995: Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages
	Subfamily SYLLINAE Grube, 1850
		Genus SYLLIS Savigny, 1818
			Syllis (Ehlersia) heterochaeta Moore, 1909
				Dorsey & Phillips, 1987: Bull. Biol. Soc. Wash. 7: 153-156, f.1
***			Syllis (Ehlersia) hyperioni ...
				Harris, 1995: Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages
			Syllis (S.) elongata Johnson, 1901
			Syllis (S.) spongiphila Verrill, 1885
			Syllis (Typosyllis) adamantea (Treadwell, 1914)
			Syllis (Typosyllis) alternata Moore, 1908
			Syllis (Typosyllis) armillaris (O.F.Müller, 1771)
			Syllis (Typosyllis) harti Berkeley & Berkeley, 1938
			Syllis (Typosyllis) hyalina Grube, 1863
			Syllis (Typosyllis) pulchra Berkeley & Berkeley, 1938
			Syllis (Typosyllis) stewarti Berkeley & Berkeley, 1942
		Genus TRYPANOSYLLIS Claparède, 1864
			Trypanosyllis (Trypanedenta) gemmipara Johnson, 1901
			Trypanosyllis (Trypanedenta) ingens Johnson, 1902
	Subfamily EUSYLLINAE Rioja, 1925
		Genus EUSYLLIS Malmgren, 1867
			Eusyllis assimilis Marenzeller, 1875
			Eusyllis blomstrandi Malmgren, 1867
			Eusyllis japonica Imajima & Hartman
			Eusyllis magnifica (Moore, 1906)
***			Eusyllis sp. P. (Harris, unpubl.)
				Harris, 1995: Re-identification of Puget Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages
		Genus ODONTOSYLLIS Claparède, 1863
			Odontosyllis fulgurans japonica Imajima
			Odontosyllis parva Berkeley, 1923
			Odontosyllis phosphorea Moore, 1909
		Genus PIONOSYLLIS Malmgren, 1867
			Pionosyllis gigantea Moore, 1908
			Pionosyllis uraga Imajima
		Genus STREPTOSYLLIS Webster & Benedict, 1884
			Streptosyllis latipalpa Banse
		Genus SYLLIDES Oersted, 1845
			Syllides japonica Imajima
			Syllides longocirrata Oersted, 1845
	Subfamily AUTOLYTINAE Langerhans, 1879
		Genus AUTOLYTUS Grube, 1850
***			Autolytus alexandri Malmgren, 1867
				Qian & Chia, 1989: Invert. Rep. Develop. 15: 49-56.
			Autolytus magnus Berkeley, 1923
			Autolytus verrilli Marenzeller, 1892
		Genus PROCEREA Ehlers, 1864
			Proceraea cornuta (Agassiz, 1862)
			Proceraea prismatica (Fabricius, 1776)
			Proceraea trilineata (Berkeley & Berkeley, 1945)

Family NEREIDIDAE Johnston, 1865
				Fitzhugh, 1987: Bull. Biol. Soc. Wash. 7: 174-183
				Glasby, 1991: Bull. Mar. Sci. 48: 559-573
	Subfamily NEREIDINAE Johnston, 1865
		Genus CERATONEREIS Kinberg, 1866
			Ceratonereis paucidentata (Moore, 1903)
		Genus CHEILONEREIS Benham, 1916
			Cheilonereis cyclurus (Harrington, 1897)
		Genus EUNEREIS Malmgren, 1865
			Eunereis wailesi Berkeley & Berkeley, 1954
		Genus NEANTHES Kinberg, 1866
			Neanthes brandti (Malmgren, 1866)
			Neanthes limnicola Johnson, 1903
			Neanthes virens (Sars, 1835)
		Genus NEREIS Linnaeus, 1758
			Nereis eakini Hartman, 1936
			Nereis grubei (Kinberg, 1866)
			Nereis neoneanthes Hartman, 1948
			Nereis pelagica Linnaeus, 1761
			Nereis procera Ehlers, 1868
			Nereis vexillosa Grube, 1851
		Genus NICON Kinberg, 1866
			[?] Nicon moniloceras (Hartman, 1940)
		Genus PERINEREIS Kinberg, 1866
			Perinereis monterea (Chamberlin, 1918)
		Genus PLATYNEREIS Kinberg, 1866
			Platynereis bicanaliculata (Baird, 1863)
	Subfamily NOTOPHYCINAE Knox & Cameron, 1970
		Genus MICRONEREIS Claparède, 1863
				Paxton, 1983: Rec. Austral. Mus. 35: 1-18.
			Micronereis nanaimoensis Berkeley & Berkeley, 1953
				Paxton, 1983: Rec. Austral. Mus. 35: 12-13, fs.1, 28-30

Family NEPHTYIDAE Grube, 1850
		Genus AGLAOPHAMUS Kinberg, 1866
			Aglaophamus rubella anops Hartman, 1950
		Genus NEPHTYS Cuvier, 1917, in Audouin & Milne-Edwards, 1833
			Nephtys assignis Hartman, 1950
			Nephtys brachycephala Moore, 1903
			Nephtys caeca (Fabricius, 1780)
			Nephtys caecoides Hartman, 1938
			Nephtys californiensis Hartman, 1938
			Nephtys ciliata (O.F.Müller, 1776)
**			Nephtys cornuta Berkeley & Berkeley, 1945
				[= N. c. franciscana Clark & Jones]
				Ellis & Ronaldson, 1988: Can. J. Zool. 66: 1898-1900
			Nephtys ferruginea Hartman, 1940
			Nephtys longosetosa Oersted, 1843
			Nephtys punctata Hartman, 1938
			Nephtys rickettsi Hartman, 1938

Family SPHAERODORIDAE Malmgren, 1867
		Genus SPHAERODOROPSIS Hartman & Fauchald, 1971
				Fauchald, 1974: J. nat. Hist. 8: 270-271
				Kudenov, 1987: Proc. Biol. Soc. Wash. 100: 917-926
			Sphaerodoropsis minuta (Webster & Benedict, 1887)
				Fauchald, 1974: J. nat. Hist. 8: 275, f.3:7
			Sphaerodoropsis sphaerulifer (Moore, 1909)
				Fauchald, 1974: J. nat. Hist. 8: 277, f.3:1-4
***			Sphaerodoropsis sp.
				Fournier & Levings, 1982: Syllogeus 40: 24
		Genus SPHAERODORUM Oersted, 1843
				Fauchald, 1974: J. nat. Hist. 8: 278
			Sphaerodorum papillifer Moore, 1909
				Fauchald, 1974: J. nat. Hist. 8: 280, f.4:6-10.

Family GLYCERIDAE Grube, 1850
		Genus GLYCERA Savigny, 1818
			Glycera americana Leidy, 1855
			Glycera convoluta Keferstein, 1862
			Glycera gigantea Quatrefages, 1865
**			Glycera nana Johnson, 1901, 1901
				[= G. capitata in B&H]
				Hilbig, 1992: Proc. Biol. Soc. Wash. 105: 716-719, f.4
			Glycera robusta Ehlers, 1868
			Glycera siphonostoma (delle Chiaje, 1825)
			Glycera tesselata Grube, 1863
		Genus HEMIPODUS Quatrefages, 1865
			Hemipodus borealis Johnson, 1901

Family GONIADIDAE Kinberg, 1866
		Genus GLYCINDE F.Müller, 1858
			Glycinde armigera Moore, 1911
			Glycinde picta Berkeley, 1927
			Glycinde polygnatha Hartman, 1950
		Genus GONIADA Audouin & Milne-Edwards, 1833
			Goniada annulata Moore, 1905
			Goniada brunnea Treadwell, 1906
			Goniada maculata Oersted, 1843

Family DORVILLEIDAE Chamberlin, 1919
				Orensanz, 1990: Antarct. Res. Ser. 52: 110-113
				Eibye-Jacobsen & Kristensen, 1994: Zool. Scr. 23: 107-131
				Alternative hypotheses on the pgylogeny of Dorvilleids, Iphitimids and Dinophilids differ only slightly, and almost axclusively in the rooting of the cladograms.
		Genus DORVILLEA Parfitt, 1866
				Wolf, 1986: Proc. Biol. Soc. Wash. 99: 627-638.
			Dorvillea (D.) moniloceras (Moore, 1909)
			Dorvillea (D.) pseudorubrovittata E.Berkeley, 1927
			Dorvillea (Schistomeringos) annulata (Moore, 1906)
			Dorvillea (Schistomeringos) japonica (Annenkova, 1937)
**			Dorvillea (Schistomeringos) longicornis (Ehlers, 101)
				Jumars, 1974: Zool. J. Linn. Soc. 54
				Fournier & Levings, 1982: Syllogeus 40: 27
				[= D. rudolphi (delle Chiaje) in B&H]
		Genus PAROUGIA Wolf, 1986
				Wolf, 1986: Proc. Biol. Soc. Wash. 99: 636-637.
*			Parougia caeca (Webster & Benedict, 1884)
				[= Dorvillea c. in B&H]
		Genus PETTIBONEIA Orensanz, 1973
***			Pettiboneia pugettensis (Armstrong & Jumars, 1978)
				Orensanz, 1990: Antarct. Res. Ser. 52: 114
				Blake & Hilbig, 1990: Bull.South. Calif. Acad. Sci.: 109-111, f.1
		Genus PROTODORVILLEA Pettibone, 1961
				Orensanz, 1990: Antarct. Res. Ser. 52: 114
			Protodorvillea gracilis (Hartman, 1938)

Family IPHITIMIDAE Fauchald, 1970
				Orensanz, 1990: Antarct. Res. Ser. 52: 116-120
		Genus OPHRYOTROCHA Claparède & Metschnikow, 1869
				Currently being reviewed by Dr. F. Pleijel (Stockholm)
			Ophryotrocha vivipara Banse
		Genus PALPIPHITIME Orensanz, 1990
				Orensanz, 1990: Antarct. Res. Ser. 52: 125
***			Palpiphitime sp.
				Mrs. Connie Retzer (Univ. of Victoria) is studying a species that may be related to this genus. Specimens originated from an abandoned salmon farm site in British Columbia (Retzer, in litt.). Related forms were informally reported for California by Montagne (1983, SCAMIT Newsletter 2(3), 9 p.).

Family DINOPHILIDAE Schultz, 1852
				Westheide, 1985, in Conway-Morris et al., eds: "The Origin and Relationships of Lower Invertebrates", Oxford Univ. Press: 310-326
				Westheide, 1990: Synopses of the British Fauna # 44.
		Genus DINOPHILUS O.Schmidt, 1848
			Dinophilus kincaidi Jones & Ferguson
		Genus TRILOBODRILUS Remane, 1925
			Trilobodrilus nipponicus Uchida & Okuda, 1943

Family LUMBRINERIDAE Malmgren, 1867
				Frame, 1992: Proc. Biol. Soc. Wash. 105: 185-218
				Orensanz, 1990: Antarct. Res. Ser. 52: 72-76
		Genus ERANNO Kinberg, 1865
*			Eranno lagunae (Fauchald, 1970)
		Genus LUMBRINERIS Blainville, 1828
				Frame, 1992: Proc. Biol. Soc. Wash. 105: 197-198
				Species in the restricted definition of this genus are likely a monophyletic group. The nomenclatorial status of the genus is uncertain because the questionable status of the type species.
			Lumbrineris californiensis Hartman, 1944
			Lumbrineris cruzensis Hartman, 1944
			Lumbrineris inflata Moore, 1911
			Lumbrineris japonica (Marenzeller, 1879)
			Lumbrineris latreilli Audouin & Milne-Edwards, 1834
			[?] Lumbrineris limicola Hartman, 1944
			Lumbrineris pallida Hartman, 1944
		Genus NINOE Kinberg, 1865
				Orensanz, 1990: Antarct. Res. Ser. 52: 98-100.
Q			Ninoe sp.
				[= N. gemmea in B&H]
				Specimens from Puget Sound labeled as such that I examined were not N. gemmea (originally described from California), but rather close to N. japonica Imajima (originally described from Japan).
		Miscellaneous lumbrinerids with simple but no composite hooded hooks
				Some species in this heterogeneous group may belong into Scoletoma Blainville (see Frame, 1992: Proc. Biol. Soc. Wash. 105: 203). Changes in nomenclature (mostly through creation of new genera) are certain to take place in the future.
			"Lumbrineris" bicirrata Treadwell, 1929
			"Lumbrineris" luti Berkeley & Berkeley, 1945
			"Lumbrineris" similabris Treadwell, 1926
			"Lumbrineris" zonata (Johnson, 1901)

Family OENONIDAE Kinberg, 1865 (= ARABELLIDAE Hartman, 1944)
				Orensanz, 1990: Antarct. Res. Ser. 52: 100-102.
		Genus ARABELLA Grube, 1850
				Colbath, 1989: Proc. Biol. Soc. Wash. 102: 283-299
Q			Arabella iricolor (Montagu, 1804)
				Originally described from the UK. Worldwide record are likely to correspond to other species; records from the Pacific NW require validation.
			[?] Arabella semimaculata (Moore, 1911)
		Genus DRILONEREIS Claparède, 1870
			Drilonereis falcata minor Hartman, 1965
			Drilonereis longa Webster, 1879
		Genus NOTOCIRRUS Schmarda, 1861
			Notocirrus californiensis Hartman, 1944

Family ONUPHIDAE Kinberg, 1865
				Paxton, 1986: Rec. Austral. Mus. 38: 1-74.
	Subfamily HYALINOECIINAE Paxton, 1986
		Genus NOTHRIA Malmgren, 1866
				Fauchald, 1982: Smith. Contr. Zool. 356: 88-89, f.26
				Kucheruk, 1985: [Explor. Fauna Seas 34(42): 83-87] (in Russian)
				Paxton, 1986: Rec. Austral. Mus. 38: 25-27, fs.15a-n
				Jirkov & Yermolaev, 1989: Zool. Zh. 58: 5-13.
Q			Nothria sp.
				[= N. conchylega in B&H]
				Worldwide records of "N. conchylega" correspond to several species (see Fauchald, 1982, for discussion). Kucheruk (1985, op.cit.) indicated occurrence of 3 species off the Pacific NW (his f.1 & 2): N. britannica, N. atlantica and N. solenotecton, the two latter presumably from deep water. Jirkov & Yermolaev (1989, op.cit.) placed N. britannica sensu Kucheruk in the synonymy of N. hyperborea. Records from the Pacific NW are in need of revision.
	Subfamily ONUPHINAE Kinberg, 1865
		Genus DIOPATRA Audouin & Milne-Edwards, 1833
				Paxton, 1986: Rec. Austral. Mus. 38: 38-41, fs.9b, 23a-l
			Diopatra ornata Moore
		Genus EPIDIOPATRA Augener, 1918
				Paxton, 1986: Rec. Austral. Mus. 38: 41, f.24a-k
Q			Epidiopatra hupferiana monroi Day
				Juveniles of Diopatra have been often reported as Epidiopatra (Paxton, 1986). Records from the Pacific NW require validation.
		Genus ONUPHIS Audouin & Milne-Edwards, 1833
				Fauchald, 1982: Smith. Contr. Zool. 356: 36-39
				Paxton, 1986: Rec. Austral. Mus. 38: 56-58, fs.3, 6-8, 10
			Onuphis elegans (Johnson, 1901)
				Fauchald, 1982: Smith. Contr. Zool. 356: 45
			Onuphis geophiliformis (Moore, 1903)
				Fauchald, 1982: Smith. Contr. Zool. 356: 47-48, f.13a, table 14
			Onuphis iridescens (Johnson, 1901)
				Fauchald, 1982: Smith. Contr. Zool. 356: 49, f.14a
		Genus MOOREONUPHIS Fauchald, 1982
				Fauchald, 1982: Smith. Contr. Zool. 356: 55-56
				Paxton, 1986: Rec. Austral. Mus. 38: 56, f.35a-j
*			Mooreonuphis stigmatis (Treadwell, 1922)
				Fauchald, 1982: Smith. Contr. Zool. 356: 

Family EUNICIDAE Savigny, 1818
		Genus EUNICE Cuvier, 1817
				Fauchald, 1992: Smith. Contr. Zool. 523, 422 p.
Q			Eunice aphroditois (Pallas, 1788)
				Fauchald, 1992, Smith.Contr.Zool.523: 62-63
				Type locality is Sri Lanka. Recorded from many tropical and warm temperate areas; possibly a species group rather than a single species. Status of form from Pacific NW needs reevaluation.
			Eunice valens (Chamberlin, 1919)
				Fauchald, 1992: Smith. Contr. Zool. 523: 332-333, ill.
				Closely relared to E. kobiensis McIntosh, 1885, of which it is considered a junior synonym by some authors (Fournier & Levings, 1982: Syllogeus 40: 24-25)

Family ORBINIIDAE Hartman, 1942
	Subfamily PROTOARICIINAE Hartman, 1957
		Genus ORBINIELLA Day, 1954
			Orbiniella nuda Hobson
		Genus PROTOARICIELLA Hartmann-Schröder, 1962
			Protoariciella oligobranchia Hobson, 1976
	Subfamily ORBINIINAE Hartman, 1942
		Genus LEITOSCOLOPLOS Day, 1977
				Mackie, 1987: Sarsia 72: 1-28
			Leitoscoloplos pugettensis (Pettibone, 1957)
				Mackie, 1987: Sarsia 72: 8-9, f.8
		Genus NAINEREIS Blainville, 1828
			Nainereis dendritica (Kinberg, 1867)
			Nainereis quadricuspida (Fabricius, 1780)
			Nainereis uncinata Hartman, 1957
		Genus PHYLO Kinberg, 1866
			Phylo felix Kinberg, 1866
		Genus SCOLOPLOS Blainville, 1828
			Scoloplos acmeceps Chamberlin, 1919
			Scoloplos armiger a. (O.F.Müller, 1776)
**Q			[?] Scoloplos armiger alaskensis (Hartman, 1948)
				[?= Leitoscoloplos panamensis in H&B]
				Mackie, 1987: Sarsia 72: 20-22, f.21

Family QUESTIDAE Hartman, 1966
				Giere & Riser, 1981: Zool. Scr. 10: 95-103
		Genus QUESTA Hartman, 1966
			Questa caudicirra Hartman, 1966

Family PARAONIDAE Cerruti, 1909
				Strelzov, 1973 (translated 1979: Polychaete Worms of the Family PARAONIDAE Cerruti, 1909, 212 pp. New Delhi: Amerind Publ.)
				Melville, 1979: Bull. Zool. Nom. 36: 114-118
		Genus ARICIDEA Webster, 1879
				Hartley, 1981: J. mar. biol. Ass. U.K. 61: 135
			Aricidea (A.) minuta Southward, 1956
Q			Aricidea neosuecica Hartman, 1965
				Status of records in the A.suecica/simplex-group requires clarification (Fournier & Levings, 1982: Syllogeus 40: 29-30)
			Aricidea (Acmira) assimilis Tebble, 1959
			Aricidea (Acmira) lopezi Berkeley & Berkeley, 1956
			Aricidea (Allia) quadrilobata Webster & Benedict, 1887
			Aricidea (Allia) ramosa Annenkova, 1934
		Genus CIRROPHORUS Ehlers, 1908
			Cirrophorus branchiatus Ehlers, 1908
		Genus LEVINSENIA Mesnil, 1897
*			Levinsenia gracilis (Tauber, 1879)
				[= Tauberia g. in H&B]
				Melville, 1979: Bull. Zool. Nom. 36: 114-118
				Hartley, 1981: J. mar. biol. Ass. U.K. 61: 135, 146
		Genus PARADONEIS Hartman, 1965
Q*			Paradoneis lyra (Southern, 1914)
				Mackie, 1991: Ophelia Suppl. 5: 148-150, fs.1-3A
				Worldwide records of this species seem to correspond to several related but distinct form; records outside the NE Atlantic require validation.
		Genus PARAONELLA Strelzov, 1973
			Paraonella platybranchia (Hartman, 1961)
			Paraonella spinifera (Hobson, 1972)

Family APISTOBRANCHIDAE Mesnil & Caullery, 1898
		Genus APISTOBRANCHUS Levinsen, 1883
			Apistobranchus ornatus Hartman, 1965
			[?] Apistobranchus tullbergi (Theel, 1879)

Family SPIONIDAE Grube, 1850
		Genus LAONICE Malmgren, 1867
Q			Laonice cirrata (M.Sars, 1850)
				Sikorsky & al., 1988: Zool. Zh. 67: 826-838
				Orrhage & Sundberg, 1990: Zool. Scr. 19: 173-178
				Several species may be recorded under this name in boreal and Arctic seas.
			Laonice pugettensis Banse & Hobson, 1968
		Genus MALACOCEROS Quatrefages, 1843
			Malacoceros (M.) fuliginosus (Claparède, 1870
			Malacoceros (Rhynchospio) glutaeus (Ehlers, 1897)
		Genus MICROSPIO Mesnil, 1896
***			Microspio sp.
				Fournier & Levings, 1982: Syllogeus 40: 30-31
		Genus PYGOSPIO Claparède, 1863
			Pygospio elegans Claparède, 1863
		Genus SCOLELEPIS Blainville, 1828
				Maciolek, 1987: Bull. Biol. Soc. Wash. 7: 16-40
			Scolelepis (S.) foliosa (Audouin & Milne-Edwards, 1833)
				Maciolek, 1987: Bull. Biol. Soc. Wash. 7: 27-30, fs.6,7
			Scolelepis squamata (Müller, 1806)
				Maciolek, 1987: Bull. Biol. Soc. Wash. 7: 30-33, f.8
		Genus SPIO Fabricius, 1785
				Maciolek, 1990: J. Nat. Hist. 24: 1109-1142
			Spio butleri Berkeley & Berkeley, 1954
			Spio cirrifera (Banse & Hobson, 1968)
			Spio filicornis (O.F.Müller, 1766)
		Genus SPIOPHANES Grube, 1860
			Spiophanes berkeleyorum Pettibone, 1962
			Spiophanes bombyx (Claparède, 1870)
			Spiophanes kroyeri Grube, 1860
		Genus STREBLOSPIO Webster, 1879
			Streblospio benedicti Webster, 1879
	Genera of the "PRIONOSPIO" group
		Genus PARAPRIONOSPIO Caullery, 1914
Q			Paraprionospio pinnata (Ehlers, 1901)
				Maciolek, 1985: Zool. J. Linn. Soc. 84: 372-374.
				Originally described from Chile. Worldwide records may correspond to several different species (i.e. Yokoyama & Tamai, 1981: Publ. Serto. Mar. Biol. Lab. 26: 303-317); records from the Pacific NW require validation.
		Genus PRIONOSPIO Malmgren, 1867
				Maciolek, 1985: Zool. J. Linn. Soc. 84: 325-383
***			Prionospio (Minuspio) multibranchiata Berkeley, 1927
				[= P. cirrifera, in part, in H&B]
				Maciolek, 1985: Zool. J. Linn. Soc. 84: 365-367, f.15.
***			Prionospio (Minuspio) lighti Maciolek, 1985
				[= P. cirrifera, in part, in H&B]
				Maciolek, 1985: Zool. J. Linn. Soc. 84: 363-364, f.14.
**			Prionospio (P.) sp.
				[= P. steenstrupi in H&B]
				Sigvaldadottir & Mackie, 1993: Sarsia 78: 203-219
				Worldwide records previously of P. steenstrupi and P. fallax are in need of revision, and probably correspond to other species in the "steenstrupi" group.
	Genera of the "POLYDORA" group
		Genus BOCCARDIA Carazzi, 1893
			Boccardia columbiana (E.Berkeley, 1927)
			Boccardia pugettensis Blake, 1979
			Boccardia polybranchia (Haswell, 1885)
				Blake, 1983: Antarct. Res. Ser. 39: 248-249
			Boccardia proboscidea Hartman, 1940
		Genus BOCCARDIELLA Blake & Kudenov, 1978
*			Boccardiella hamata (Webster, 1879)
				[= Polydora (Boccardia) h. in H&B]
				Blake & Kudenov, 1978: Mem. Nat. Mus. Victoria 39: 264-265
				Radashevsky, 1993: Publ. Seto Mar. Biol. Lab. 36: 4-7, fs.2-3
		Genus POLYDORA Bosc, 1802
			Polydora alloporis Light, 1970
			Polydora brachycephala Hartman, 1936
			Polydora cardalia E.Berkeley, 1927
				Radashevsky, 1993: Publ. Seto Mar. Biol. Lab. 36: 23-28, fs.11-12
***			Polydora caulleryi Mesnil, 1897
				Fournier & Levings, 1982: Syllogeus 40: 31
			Polydora commensalis Andrews, 1891
				Radashevsky, 1993: Publ. Seto Mar. Biol. Lab. 36: 12-15, f.6
**			Polydora cornuta Bosc, 1802
				[= P. ligni in H&B]
				Blake & Maciolek, 1987: Bull. Biol. Soc. Wash. 7: 11-15
			Polydora giardi Mesnil, 1896
			Polydora pygidialis Blake & Woodwick, 1972
			Polydora quadrilobata Jacobi, 1883
				Radashevsky, 1993: Publ. Seto Mar. Biol. Lab. 36: 18-21, f.9
			Polydora socialis (Schmarda, 1861)
			Polydora spongicola Berkeley & Berkeley, 1950
				Radashevsky, 1993: Publ. Seto Mar. Biol. Lab. 36: 21-23, f.10
		Genus PSEUDOPOLYDORA Czerniavsky, 1881
			Pseudopolydora kempi japonica Imajima & Hartman, 1964

Family TROCHOCHAETIDAE Pettibone, 1963
		Genus TROCHOCHAETA Levinsen, 1883
			Trochochaeta multisetosa (Oersted, 1844)

Family MAGELONIDAE Cunningham & Ramage, 1888
		Genus MAGELONA F.Müller, 1858
			Magelona berkeleyi Jones, 1971
			Magelona hobsonae Jones, 1978
			Magelona longicornis Johnson, 1901
			Magelona sacculata Hartman, 1961

Family CHAETOPTERIDAE Malmgren, 1867
		Genus CHAETOPTERUS Cuvier, 1827
Q			Chaetopterus variopedatus (Renier, 1804)
				Most world-wide citations of this species correspond to other genera and species (Mrs. Mary Petersen, pers. comm.). Records from the Pacific NW require validation.
		Genus MESOCHAETOPTERUS Potts, 1914
			Mesochaetopterus taylori Potts, 1914
		Genus PHYLLOCHAETOPTERUS Grube, 1863
			Phyllochaetopterus claparedii McIntosh, 1885
			Phyllochaetopterus prolificus Potts, 1914
		Genus SPIOCHAETOPTERUS Sars, 1853
			Spiochaetopterus costarum (Claparède, 1870)

Family COSSURIDAE Day, 1963
				Family being reviewed for the NE Pacific by Dr. Judith Fournier (Canadian Museum of Nature, Ottawa). At least two species will be added.
		Genus COSSURA Webster & Benedict, 1887
				Fournier & Petersen, 1991: Ophelia 5 Suppl.: 63-80
***			Cossura modica Fauchald & Hancock, 1981
				Fournier & Levings, 1982: Syllogeus 40: 35
**			Cossura pygodactylata Jones, 1956
				Fournier & Petersen, 1991: Ophelia 5 Suppl.: 76-77.
				[= Cossura soyeri in B&H]
			Cossura spp.

Family CIRRATULIDAE Carus, 1863
	Genera with one pair of tentacular palps
			Anonymous, 1995: Notes on Aphelochaeta and Monticellina. Pacific NW Polychaete Worksop, informal document, 9 pages.
		Genus APHELOCHAETA Blake, 1991
				Blake, 1991: Ophelia Suppl. 5: 28
***			[?] Aphelochaeta monilaris (Hartman, 1960)
				Fournier & Levings, 1982: Syllogeus 40: 33-34 (as Tharyx m.)
*			Aphelochaeta multifilis (Moore, 1909)
*			Aphelochaeta parva (E.Berkeley, 1929)
*			Aphelochaeta secunda (Banse & Hobson, 1968)
*			Aphelochaeta serratiseta (Banse & Hobson, 1968)
		Genus CAULLERIELLA Chamberlin, 1919
			[?] Caulleriella alata (Southern, 1914)
		Genus CHAETOZONE Malmgren, 1867
			Chaetozone acuta Banse & Hobson, 1968
***			Chaetozone berkeleyorum Banse & Hobson, 1968
				Fournier & Levings, 1982: Syllogeus 40.
				Considered a synonym of Cirratulus cirratus in H&B, following a personal communication from Dr. P. Jumars, from the U. of Washington).
			Chaetozone setosa Malmgren, 1867
			[?] Chaetozone spinosa Moore, 1903
		Genus DODECACERIA Oersted, 1843
Q			Dodecaceria concharum Oersted, 1843
				George & Petersen, 1991: Ophelia Suppl.5: 89-100
				European records of this species correspond to two different forms: D. concharum and D. ater (Quatrefages, 1866). Records from world-wide areas require validation.
			Dodecaceria fewkesi Berkeley & Berkeley, 1954
		Genus MONTICELLINA Laubier, 1961
				Blake, 1991: Ophelia Suppl. 5: 24
***			Monticellina tesselata (Hartman, 1960)
				Fournier & Levings, 1982: Syllogeus 40 (as Tharyx t.)
	Genera with clusters of tentacular palps
		Genus CIRRATULUS Lamarck, 1801, in Audouin & Milne-Edwards, 1833
			Cirratulus cirratus (O.F.Müller, 1776)
			Cirratulus spectabilis (Kinberg, 1866)

Family CTENODRILIDAE Kennel, 1882
				Petersen & George, 1991: Ophelia Suppl. 5: 185-208
	Subfamily CTENODRILINAE Kennel, 1982
		Genus CTENODRILUS Claparède, 1863
			Ctenodrilus serratus (Schmidt, 1857)
	Subfamily RAPHIDRILINAE Hartmann-Schröder, 1971
		Genus RAPHIDRILUS Monticelli, 1910
***			Raphidrilus sp.
				Qian & Chia, 1989: Can. J. Zool. 67: 2345-2351 (as R. nemasoma)
				Petersen & George, 1991: Ophelia Suppl. 5: 203-206

Family FLABELLIGERIDAE Saint-Joseph, 1894
		Genus BRADA Stimpson, 1854
			Brada sachalina Annenkova, 1922
			Brada villosa (Rathke, 1843)
		Genus FLABELLIDERMA Hartman, 1969
***			Flabelliderma sp.
				Photographs of a live specimen collected in the lower rocky intertidal zone (among barnacles) at Bamfield (BC) were shown to me by Dr. Greg Jensen (University of Washington) in 1991. They did not seem to correspond to any described species.
		Genus FLABELLIGERA M.Sars, 1829
			Flabelligera affinis M.Sars, 1829
		Genus PHERUSA Oken, 1807, in Stop-Bowitz, 1948
			Pherusa negligens (Berkeley & Berkeley, 1950)
			Pherusa plumosa (O.F.Müller, 1776)
		Genus PIROMIS Kinberg, 1867
			Piromis eruca (Claparède, 1870)

Family ACROCIRRIDAE Banse, 1969
		Genus ACROCIRRUS Grube, 1872
			Acrocirrus columbianus Banse, 1969
			Acrocirrus occipitalis Banse, 1969
		Genus MACROCHAETA Grube, 1850
			Macrochaeta pege Banse, 1969

Family SCALIBREGMIDAE Malmgren, 1867
		Genus ASCLEROCHEILUS Ashworth, 1901
			Asclerocheilus beringianus Ushakov, 1955
		Genus HYBOSCOLEX Schmarda, 1861
			Hyboscolex pacificus (Moore, 1909)
		Genus SCALIBREGMA Rathke, 1843
				Mackie, 1991: Bull. Mar. Sci. 48: 268-276.
			Scalibregma inflatum Rathke, 1843
				Mackie, 1991: Bull. Mar. Sci. 48: 268-271, fs.1-10

Family OPHELIIDAE Malmgren, 1867
	Subfamily OPHELIINAE Malmgren, 1867
		Genus EUZONUS Grube, 1866
			Euzonus mucronata (Treadwell, 1914)
			Euzonus williamsi (Hartman, 1938)
		Genus OPHELIA Savigny, 1818
				Bellan & Dauvin, 1991: Bull. Mar. Sci. 48: 544-558.
			Ophelia limacina (Rathke, 1843)
	Subfamily OPHELININAE Hartmann-Schröder, 1971
		Genus ARMANDIA Filippi, 1861
			Armandia brevis (Moore, 1906)
		Genus OPHELINA Oersted, 1843
			Ophelina acuminata (Oersted, 1843)
			Ophelina breviata (Ehlers, 1913)
	Subfamily TRAVISIINAE Hartmann-Schröder, 1971
				Dauvin & Bellan, 1994: Mém.Mus.natn.Hist.nat.162: 169-184
		Genus TRAVISIA Johnston, 1840
			Travisia brevis Moore, 1923
			Travisia forbesii Johnston, 1840
			Travisia japonica Fujiwara, 1933
			Travisia pupa Moore, 1906

Family STERNASPIDAE Carus, 1863
		Genus STERNASPIS Otto, 1821
			Sternaspis scutata (Renier, 1807)

Family CAPITELLIDAE Grube, 1862
		Genus BARANTOLLA Southern, 1921
			Barantolla americana Hartman, 1969
		Genus CAPITELLA Blainville, 1828
				Eckelbarger & Grassle, 1987: Bull. Biol. Soc. Wash. 7: 62-76
				Grassle & al., 1987: Bull. Biol. Soc. Wash. 7: 77-88.
				Warren, 1991: Ophelia Suppl. 5: 275-282
**			Capitella "capitata" hyperspecies
				World-wide records include several co-occurring, unnamed sibling species (see references listed above).
			Capitella dizonata Johnson, 1901
		Genus DECAMASTUS Hartman, 1969
			Decamastus gracilis Hartman, 1969
		Genus HETEROMASTUS Eisig, 1887
			Heteromastus filiformis (Claparède, 1864)
			Heteromastus filobranchus Berkeley & Berkeley, 1932
		Genus MEDIOMASTUS Hartman, 1944
				Warren et al., 1994: Rec. Austral. Mus. 46: 227-256
			Mediomastus ambiseta (Hartman, 1947)
				Warren et al., 1994: Rec. Austral. Mus. 46: 234-236, fs.2a, 5d-f, 6a-f, 12, Tables 1 & 2
**			Mediomastus californiensis Hartman, 1944
				Warren et al., 1994: Rec. Austral. Mus. 46: 239-241, fs. 2b, 7c-d, 9a-d, 12; Tables 1 & 2
				[Records of M. capensis Day, 1961 from the Pacific NW correspond to this species]
		Genus NOTOMASTUS Sars, 1851
			Notomastus giganteus Moore, 1906
			Notomastus lineatus Claparède, 1870
			Notomastus tenuis Moore, 1909
			Notomastus variegatus Berkeley & Berkeley, 1950

Family PARERGODRILIDAE Reisinger, 1925
				Westheide, 1990: Synopses of the British Fauna # 44.
		Genus STYGOCAPITELLA Knollner, 1934
			Stygocapitella subterranea Knollner, 1934
				Riser, 1980: Wasmann J. Biol. 38: 10-17
				Westheide, 1990: Synopses of the British Fauna 44: 138-139, f.50

Family ARENICOLIDAE Johnston, 1846
		Genus ABARENICOLA Wells, 1959
			Abarenicola claparedi oceanica Healy & Wells, 1959
			Abarenicola claparedi vagabunda Healy & Wells, 1959
			Abarenicola pacifica Healy & Wells, 1959
		Genus ARENICOLA Lamarck, 1801, in Audouin & Milne-Edwards, 1833
			Arenicola marina Linnaeus, 1758
		Genus BRANCHIOMALDANE Langerhans, 1881
**			Branchiomaldane simplex Berkeley & Berkeley, 1932
				[= B. vincenti in H&B]
				Fournier & Barrie, 1987: Bull.Biol.Soc. Wash. 7: 101-103, f.2a-f

Family MALDANIDAE Malmgren, 1867
	Subfamily CLYMENURINAE Imajima & Shiraki, 1982
		Genus CLYMENURA Verrill, 1900
			Clymenura (Cephalata) columbiana (Berkeley, 1929)
***			Clymenura (Cephalata) gracilis Hartman, 1969
				Harris & Green, 1995: Checklist of West Coast MALDANIDAE. Pacific NW Polychaete Workshop, informal document, 10 pages.
	Subfamily EUCLYMENINAE Arwidsson, 1907
		Genus AXIOTHELLA Verrill, 1900
Q			Axiothella rubrocincta (Johnson, 1901)
				Wilson, 1983: Mar. Biol. 76: 297-300
				Seems to constitute a sibling species complex.
		Genus CLYMENELLA Verrill, 1873
			Clymenella torquata (Leidy, 1855)
		Genus EUCLYMENE Verrill, 1900
			Euclymene zonalis (Verrill, 1874)
			Euclymene cf. zonalis (Verrill, 1874)
			Euclymene sp. A
				Harris, 1995: Re-identification of Puger Sound specimens. Pacific NW Polychaete Workshop, informal document, 5 pages.
		Genus ISOCIRRUS Arwidsson, 1907
			Isocirrus longiceps (Moore, 1923)
		Genus MACROCLYMENE Verrill, 1900
			Macroclymene sp.
		Genus MALDANELLA McIntosh, 1885
			Maldanella harai (Izuka, 1902)
			Maldanella robusta Moore, 1906
		Genus PRAXILLELA Verrill, 1881
				Harris, 1995: Staining patterns of selected Praxillella species. Pacific NW Polychaete Workshop, informal document, 2 pages.
			Praxillela affinis (Sars, 1872)
			Praxillela gracilis (Sars, 1861)
			Praxillela pacifica Berkeley, 1929
				[= P. affinis p. in H&B]
				Harris & Green, 1995: Checklist of West Coast MALDANIDAE. Pacific NW Polychaete Workshop, informal document, 10 pages.
			Praxillela praetermissa (Malmgren, 1866)
		Genus PRAXILLURA Verrill, 1880
			Praxillura maculata Moore, 1923
	Subfamily LUMBRICLYMENINAE Arwidsson, 1907
		Genus NOTOPROCTUS Arwidsson, 1907
			Notoproctus pacificus (Moore, 1906)
	Subfamily MALDANINAE Arwidsson, 1907
				Light, 1991: Ophelia Suppl. 5: 133-146
		Genus CHIRIMIA Light, 1991
				Light, 1991: Ophelia Suppl. 5: 136-139
*			Chirimia biceps b. (Sars, 1861)
				[= Asychis b. & A. lacera in H&B]
				Asychis b.: Harris & Green, 1995: Checklist of West Coast MALDANIDAE. Pacific NW Polychaete Workshop, informal document, 10 pages.
			Chirimia similis (Moore, 1906)
		Genus MALDANE Grube, 1840
				Light, 1991: Ophelia Suppl. 5: 136
Q			Maldane glebifex Grube, 1860
				Pacific records of this species are most likely misidentified M. sarsi; K.Green in Anonymous, 1995: Northwest Pacific Maldanids. Pacific NW Polychaete Workshop, informal document, 3 pages.
			Maldane sarsi Malmgren, 1865
		Genus METASYCHIS Light, 1991
				Light, 1991: Ophelia Suppl. 5: 141
*			Metasychis disparidentus (Moore, 1904)
				[= Asychis d. in H&B]
	Subfamily NICOMACHINAE Arwidsson, 1907
		Genus MICROMALDANE Mesnil, 1897
			Micromaldane ornithochaeta Mesnil, 1897
		Genus NICOMACHE Malmgren, 1865
			Nicomache lumbricalis (Fabricius, 1780)
			Nicomache personata Johnson, 1901
		Genus PETALOPROCTUS Quatrefages, 1865
			Petaloproctus borealis Arwidsson, 1907
				[as P. tenuis b. in H&B]
				Harris & Green, 1995: Checklist of West Coast MALDANIDAE. Pacific NW Polychaete Workshop, informal document, 10 pages.
			Petaloproctus tenuis (Theel, 1879)
	Subfamily RHODININAE Arwidsson, 1907
			Rhodine bitorquata Moore, 1923

Family OWENIIDAE Rioja, 1917
				Nilsen & Holthe, 1985: Sarsia 70: 17-32
		Genus MYRIOCHELE Malmgren, 1867
				Opinion 1636. 1991. Bull. Zool. Nom. 48: 164-165
			Myriochele oculata Zachs, 1923
				Opinion 1636. 1991. Bull. Zool. Nom. 48: 164-165
		Genus OWENIA delle Chiaje, 1842
			Owenia fusiformis delle Chiaje, 1842
				Dauvin & Thiébaut, 1994: Mém.Mus.natn.Hist.nat.162: 383-404

Family SABELLARIIDAE Johnston, 1865
				Kirtley, 1994: A review and taxonomic revision of the family SABELLARIIDAE Johnston, 1865, Sabecon Press, Sci. Ser. No. 1, 223 pp.
		Genus IDANTHYRSUS Kinberg, 1867
				Kirtley, 1994, op.cit.: 84-86
**			Idanthyrsus saxicavus (Baird, 1963)
				[= I. armatus + I. ornamentatus in H&B]
				Kirtley, 1994, op.cit.: 108-113, fs. 6.15.1-7
		Genus NEOSABELLARIA Kirtley, 1994
				Kirtley, 1994, op.cit.: 16-17
*			Neosabellaria cementarium (Moore, 1906)
				[= Sabellaria c. in H&B]
				Kirtley, 1994, op.cit.: 18-22

Family AMPHICTENIDAE Malmgren, 1867
				Holthe, 1986: Marine Invert. Scandin. 7
		Genus AMPHICTENE Savigny, 1818
			Amphictene moorei Annenkova, 1929
		Genus CISTENIDES Malmgren, 1866
			Cistenides granulata (Linnaeus, 1767)
				Holthe, 1986: Marine Invert. Scandin. 7: 24-25, f.5.
		Genus PECTINARIA Savigny, 1818
			Pectinaria californiensis Hartman, 1941

Family AMPHARETIDAE Malmgren, 1867
				Holthe, 1986: Marine Invert. Scandin. 7
	Subfamily AMPHARETINAE Chamberlin, 1919
		Genus AMAGE Malmgren, 1866
			Amage anops (Johnson, 1901)
		Genus AMPHARETE Malmgren, 1866
			Ampharete acutifrons (Grube, 1860)
				Holthe, 1986: Marine Invert. Scandin. 7: 33-34, f.8
			Ampharete finmarchica (M.Sars, 1866)
				Holthe, 1986: Marine Invert. Scandin. 7: 38-40, f.11
			Ampharete goesi brazhnikovi Annenkova, 1929
			Ampharete goesi g. Malmgren, 1866
				Holthe, 1986: Marine Invert. Scandin. 7: 40-41, f.12
			Ampharete labrops Hartman, 1961
		Genus AMPHICTEIS Grube, 1850
			Amphicteis mucronata Moore, 1923
			Amphicteis scaphobranchiata Moore, 1906
		Genus ANOBOTHRUS Levinsen, 1884
			Anobothrus gracilis (Malmgren, 1866)
				Holthe, 1986: Marine Invert. Scandin. 7: 50-51, f.18
		Genus ASABELLIDES Annenkova, 1929
			Asabellides lineata (Berkeley & Berkeley, 1943)
			Asabellides sibirica (Wirén, 1883)
		Genus HOBSONIA Banse, 1979
			Hobsonia florida (Hartman, 1951)
		Genus LYSIPPE Malmgren, 1866
			Lysippe labiata Malmgren, 1866
				Holthe, 1986: Marine Invert. Scandin. 7: 52-54, f.19
		Genus MOORESAMYTHA Williams, 1987
*			Mooresamytha bioculata (Moore, 1906)
				[= Amphisamytha b. in H&B]
				Williams, 1987: Bull. Biol. Soc. Wash. 7: 256-257, f.4A,B
		Genus SAMYTHA Malmgren, 1866
			Samytha californiensis Hartman, 1969
		Genus SCHISTOCOMUS Chamberlin, 1919
			Schistocomus hiltoni Chamberlin, 1919
		Genus SOSANOPSIS Hessle, 1917
			Sosanopsis hesslei Banse, 1979
	Subfamily MELINNINAE Chamberlin, 1919
		Genus MELINNA Malmgren, 1866
			Melinna cristata (Sars, 1851)
				Holthe, 1986: Marine Invert. Scandin. 7: 81-83, f.34
			Melinna elisabethae McIntosh, 1922

Family TEREBELLIDAE Malmgren, 1866
				Holthe, 1986: Marine Invert. Scandin. 7
	Subfamily POLYCIRRINAE Malmgren, 1866
		Genus AMAEANA Hartman, 1959
			Amaeana occidentalis (Hartman, 1944)
		Genus LYSILLA Malmgren, 1866
			Lysilla loveni Malmgren, 1866
				Holthe, 1986: Marine Invert. Scandin. 7: 160-162, f.75
		Genus POLYCIRRUS Grube, 1850
				Lovell, 1995: Selected Polycirrus from Puget Sound, informal document, 2 pages, NW Polychaete Workshop.
			Polycirrus californicus Moore, 1909
			Polycirrus nr. californicus
			Polycirrus sp. I
			Polycirrus sp. II
			Polycirrus sp. III
			Polycirrus sp. IV
			Polycirrus sp. V
			Polycirrus sp. A
			Polycirrus sp. B
	Subfamily THELEPODINAE Hessle, 1917
				Hutchings & Glasby, 1987: Bull. Biol. Soc. Wash. 7: 217-250
		Genus STREBLOSOMA Sars, 1872
			Streblosoma bairdi (Malmgren, 1866)
				Holthe, 1986: Marine Invert. Scandin. 7: 142-143, f.64
		Genus THELEPUS Leuckart, 1849
			Thelepus cincinnatus (Fabricius, 1780)
				Holthe, 1986: Marine Invert. Scandin. 7: 140-142, f.63
			Thelepus crispus Johnson, 1901
***			Thelepus hamatus Moore, 1905
				[Included in H&B's keys]
				Fournier & Levings, 1982: Syllogeus 40.
			? Thelepus japonicus Marenzeller, 1884
			Thelepus setosus (Quatrefages, 1865)
	Subfamily ARTACAMINAE Malmgren, 1866
		Genus ARTACAMA Malmgren, 1866
			Artacama conifera Moore, 1905
	Subfamily AMPHITRITINAE Malmgren, 1866
		Genus AMPHITRITE O.F.Müller, 1771
			Amphitrite cirrata O.F.Müller, 1771 in 1776
				Holthe, 1986: Marine Invert. Scandin. 7: 96-98, f.40
		Genus BETAPISTA Banse, 1980
			Betapista dekkerae Banse, 1980
		Genus EUPOLYMNIA Verrill, 1900
			Eupolymnia heterobranchia (Johnson, 1901)
		Genus LANASSA Malmgren, 1866
			Lanassa venusta (Malm, 1874)
				Holthe, 1986: Marine Invert. Scandin. 7: 128-129, f.57
		Genus LAPHANIA Malmgren, 1866
			Laphania boecki Malmgren, 1866
				Holthe, 1986: Marine Invert. Scandin. 7: 130-131, f.58
		Genus NEOAMPHITRITE Hessle, 1917
			Neoamphitrite edwardsi (Quatrefages, 1865)
			Neoamphitrite robusta (Johnson, 1901)
		Genus NEOLEPREA Hessle, 1917
			Neoleprea japonica Hessle, 1917
			Neoleprea spiralis (Johnson, 1901)
	Genus NICOLEA Malmgren, 1866
			Nicolea zostericola (Oersted, 1844 in Grube, 1860)
				Holthe, 1986: Marine Invert. Scandin. 7: 124-126, f.55
		Genus PISTA Malmgren, 1866
				Saphronova, 1988: Zool. Zh. 67: 888-897.
				Saphronova, 1991: Ophelia Suppl. 5: 243-246
				Smith, 1992: J. Morphol. 213: 365-393
				Lovell, 1995: Selected Pista from Puget Sound. Pacific NW Polychaete Workshop; informal document, 2 pages.
				Genus needs review; species constitute a heterogeneous assemblage. 
***			Pista bansei Saphronova, 1988
				Saphronova, 1988: Zool. Zh. 67: 892-894, fs.
				May be a synonym of Betapista dekkerae (Harris & Lovell, pers. comm.)
			Pista brevibranchiata Moore, 1923
Q			Pista cristata (O.F.Müller, 1776)
				Holthe, 1986: Marine Invert. Scandin. 7: 112-114, f.49
				Records of this species may correspond to P. bansei, P. wui, or other members of the "cristata" group (Saphronova, 1988, op.cit.)
Q			"Pista" elongata Moore, 1909
				Does not fit the generic diagnosis (Smith, 1992: J. Morphol. 213)
*			Pista estevanica (Berkeley & Berkeley, 1942)
				[= Scionella e. in H&B]
				Saphronova, 1991: Ophelia Suppl. 5: 244-245, f.4
			Pista moorei Berkeley & Berkeley, 1942
Q			"Pista" pacifica Berkeley & Berkeley, 1942
				Does not fit the generic diagnosis (Smith, 1992, op.cit.)
			Pista wui Saphronova, 1988
				Saphronova, 1988: Zool. Zh. 67: 894-895, fs.
		Genus PROCLEA Saint-Joseph, 1894
			Proclea graffi (Langerhans, 1884)
				Holthe, 1986: Marine Invert. Scandin. 7: 132-134, f.59
		Genus SCIONELLA Moore, 1903
				Saphronova, 1991: Ophelia Suppl. 5: 239-247
			Scionella japonica Moore, 1903
				Saphronova, 1991: Ophelia Suppl. 5: 240-241
	Genus TEREBELLA Linnaeus, 1767
			Terebella ehrenbergi Grube, 1870

Family TRICHOBRANCHIDAE Malmgren, 1866
				Holthe, 1986: Marine Invert. Scandin. 7
		Genus NOVOBRANCHUS Berkeley & Berkeley, 1954
			Novobranchus pacificus Berkeley & Berkeley, 1954
		Genus TEREBELLIDES Sars, 1835
				Lovell, 1995: Selected Terebellides from Puget Sound. Pacific NW Polychaete Workshop, informal document, one page (table).
***			Terebellides californica Williams, 1984
***			Terebellides nr. horikoshii
***			Terebellides kobei Hessle, 1917
***			Terebellides nr. lineata
***			Terebellides reishi Williams, 1984
			Terebellides stroemi M.Sars , 1835
				Williams, 1984: Proc. 1st Intl. Polych. Conf.: 118-142.
				Holthe, 1986: Marine Invert. Scandin. 7: 170-172, f.80
				Pre-1995 records correspond in part to the other species listed.
		Genus TRICHOBRANCHUS Malmgren, 1866
			Trichobranchus glacialis Malmgren, 1866
				Holthe, 1986: Marine Invert. Scandin. 7: 164-165, f.77

Family SABELLIDAE Johnston, 1846
				Fitzhugh, 1989: Bull. Am. Mus. Nat. Hist. 192, 104 p.
				Fitzhugh, 1991: Zool J. Linn. Soc. 102: 305-332.
				Smith, 1991: Ophelia Suppl. 5: 249-250
				Distribution of genera among families, below, follows Fitzhugh (1989, 1991).
	Subfamily FABRICIINAE Rioja, 1923
		Genus FABRICIA Blainville, 1828
			Fabricia oregonica Banse, 1979
			Fabricia sabella (Ehrenberg, 1837)
		Genus FABRICIOLA Friedrich, 1939
				Fitzhugh, 1990: Zool. Scr. 19: 153-164
			Fabriciola berkeleyi Banse, 1956
				Fitzhugh, 1992: Pac. Sci. 46: 68-76.
		Genus MANAYUNKIA Leidy, 1858
			Manayunkia aestuarina (Bourne)
	Subfamily SABELLINAE Johnston, 1846
	   Genera traditionally placed in the FABRICIINAE
		Genus CHONE Kröyer, 1856
			Chone aurantiaca (Johnson, 1901)
			Chone duneri Malmgren, 1867
				Giangrande, 1992: Boll. Zool. 59: 525, fs.
			Chone ecaudata (Moore, 1923)
			Chone infundibiliformis Kröyer, 1856
			Chone magna (Moore, 1923)
		Genus EUCHONE Malmgren, 1866
			Euchone analis (Kröyer, 1856)
			Euchone incolor Hartman, 1965
		Genus JASMINEIRA Langerhans, 1880
			Jasmineira pacifica Annenkova, 1937
		Genus ORIOPSIS Caullery & Mesnil, 1896
			Oriopsis gracilis Hartman, 1969
			Oriopsis minuta (Berkeley & Berkeley, 1932)
				Wu, 1984: Proc. 1st Intl. Polych. Conf.: 413-417
	   Genera of SABELLINAE in the traditional definition
		Genus DEMONAX Kinberg, 1867
				Knight-Jones, 1983: Zool. J. Linn. Soc. 79: 254-255.
				Perkins, 1984: Proc. Biol. Soc. Wash. 97: 292-296.
			Demonax medius (Bush, 1905)
				Perkins, 1984: Proc. Biol. Soc. Wash. 97: 308-313, fs.11-14
**			Demonax rugosus (Moore, 1904)
				[= Sabella (D.) media, in part, in H&B]
				Perkins, 1984: Proc. Biol. Soc. Wash. 97: 304-307, fs.9,10
		Genus EUDISTYLIA Bush, 1905
			Eudistylia catharinae Banse, 1979
			Eudistylia polymorpha (Johnson, 1901)
			Eudistylia vancouveri (Kinberg, 1867)
		Genus LAONOME Malmgren, 1866
			Laonome kroyeri Malmgren, 1866
		Genus MEGALOMMA Johansson, 1927
			Megalomma splendida (Moore, 1905)
		Genus POTAMILLA Malmgren, 1866
			Potamilla neglecta (Sars, 1851)
			Potamilla intermedia (Moore, 1905)
			Potamilla myriops Marenzeller, 1884
			Potamilla occelata Moore, 1905
		Genus SABELLASTARTE Kröyer, 1856
			Sabellastarte sp.
		Genus SCHIZOBRANCHIA Bush, 1905
			Schizobranchia insignis Bush, 1904
		Sabellinae of uncertain generic possition
Q			Distylia volutacornis pacifica Berkeley & Berkeley, 1954
				[= Demonax pacifica in H&B]
				Not a species of Demonax; type material, in poor condition, can not be identified at the generic level (Perkins, 1984: Proc. Biol. Soc. Wash. 97: 294)
	   Genera tratitionally placed in the MYXICOLINAE Rioja, 1923
		Genus MYXICOLA Koch in Renier, 1847
			Myxicola aesthetica (Claparède, 1870)
			Myxicola infundibulum (Renier, 1804)

Family SERPULIDAE Savigny, 1818
				Uchida, 1978: Bull. Mar. Park Res. Stn.,# 2
				ten Hove, 1984: Proc. 1st Intl. Polych. Conf.: 181-196.
				Smith, 1991: Ophelia Suppl. 5: 249-250
	Subfamily PROTULINAE Uchida, 1978
		Genus APOMATOPSIS Saint-Joseph, 1894
*			Apomatopsis geniculata (Moore & Bush, 1904)
				[= Apomatus g. in H&B]
				Uchida, 1978: Bull. Mar. Park Res. Stn., 2: 78
		Genus APOMATUS Philippi, 1844
			Apomatus timmsii Pixell, 1912
		Genus PROTULA Saint-Joseph, 1894
			Protula pacifica Pixell, 1912
	Subfamily FILOGRANINAE Rioja, 1923
		Genus SALMACINA Claparède, 1870
			Salmacina tribranchiata (Moore, 1923)
	Subfamily SERPULINAE Savigny, 1818
		Genus CRUCIGERA Benedict, 1887
				ten Hove & Jansen-Jacobs, 1984: Proc. 1st Intl. Polych. Conf.: 143-180.
			Crucigera irregularis Bush, 1905
				ten Hove & Jansen-Jacobs, 1984: Proc. 1st Intl. Polych. Conf.: 170-176, figs.
				Kupriyanova & Rzhavsky, 1993: Ophelia 38: 51-52
			Crucigera zygophora (Johnson, 1901)
				ten Hove & Jansen-Jacobs, 1984: Proc. 1st Intl. Polych. Conf.: 165-170, figs.
				Kupriyanova & Rzhavsky, 1993: Ophelia 38: 52-53
		Genus SERPULA Linnaeus, 1767
**			Serpula columbiana Johnson, 1901
				Kupriyanova & Rzhavsky, 1993: Ophelia 38: 48-51, f.1
				Kupriyanova, 1995: Am. Zool.  ...: 114A(ref. 455), abstract only
				[= S. vermicularis in H&B]
	Subfamily HYALOPOMATINAE Uchida, 1978
		Genus PSEUDOCHITINOPOMA Zibrowius, 1969
			Pseudochitinopoma occidentalis (Bush, 1904)

Family SPIRORBIDAE
				Hess, 1993: Am. Nat. 141: 577-596
	Subfamily CIRCEINAE Knigh-Jones, 1978
		Genus CIRCEIS Saint-Joseph, 1894
***			Circeis armoricana Saint-Josep, 1894
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 427-429
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: p.193, f.2
			Circeis spirillum (Linnaeus, 1758)
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 429.
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: p.193, f.2
		Genus PARADEXIOSPIRA Caullery & Mesnil, 1897
*			Paradexiospira (P.) violacea (Levinsen, 1883)
				[= Spirorbis (Prodexiospira) v. in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 429-430.
**			Paradexiospira (Spirorbides) vitrea (Fabricius, 1780)
				[= Spirorbis (P.) nakamurai and S. (Eulaeospira) variabilis in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 430.
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: p.192, f.2
	Subfamily JANUINAE Knight-Jones, 1978
		GHenus JANUA Saint-Joseph, 1894
**			Janua pagenstecheri (Quatrefages, 1865)
				[= Spirorbis (Janua) pusilloides in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 432.
	Subfamily SPIRORBINAE
		Genus SPIRORBIS Daudin, 1800
				Records of "Spirorbis borealis" from the Pacific NW(including H&B) seem to correspond to species of Pileolaria (Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 423, bottom).
***			Spirorbis bifurcatus Knight-Jones, 1978
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 426.
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: f.1
	Subfamily PILEOLARIINAE Knight-Jones, 1978
		Genus BUSHIELLA Knight-Jones, 1973
				Knight-Jones, 1984: Zool. J. Linn. Soc. 80: 112
				Rzhavsky, 1993: Oikos 38: 89-96
***			Bushiella abnormis (Bush, 1904)
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 446-447 (as Sinistrella a.)
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: f.3
**			Bushiella verruca (Fabricius, 1780)
				[= Spirorbis (Leodora) validus in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 446 (as Sinistrella verruca)
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: f.3
		Genus JUGARIA Knight-Jones, 1978
				Knight-Jones, 1984: Zool. J. Linn. Soc. 80: 112
				Rzhavsky, 1993: Oikos 38: 89-96
***			Jugaria similis (Bush, 1904)
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 445)
**			Jugaria quadrangularis (Stimpson, 1854)
				[= Spirorbis (Pileolaria) granulatus in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 443-445
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: p.193, f.3
		Genus PILEOLARIA Claparède, 1868
				Knight-Jones, 1984: Zool. J. Linn. Soc. 80: 111
**			Pileolaria (Nidularia) dalestraughanae Vine, 1972
				[= Spirorbis (Pileolaria) langerhansi in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 442
***			Pileolaria (P.) berkeleyana (Rioja, 1942)
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 437
				Hess, 1993 Am. Nat. 141: 577-596
**			Pileolaria (Simplicaria) potswaldi Knight-Jones, 1978
				[= Spirorbis (Pileolaria) moerchi in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 441-442.
		Genus PROTLEODORA Pillai, 1970
				Knight-Jones, 1984: Zool. J. Linn. Soc. 80: 114
**			Protoleodora asperata (Bush, 1904)
				[= Spirorbis (Romanchella) medius in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 447-448 (as Sinistrella media)
				Knight-Jones & al., 1991: Bull. Mar. Sci. 48: p.194, f.3
	Subfamily ROMANCHELLINAE Knight-Jones, 1978
		Genus PROTOLAEOSPIRA Pixell, 1912
**			Protolaeospira eximia (Bush, 1904)
				[= Spirorbis (Protolaeospira) ambilateralis and S. (P.) racemosa in H&B]
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 449-450
***			Protolaeospira capensis Day, 1961
				Knight-Jones & Knight-Jones, 1979: J. Zool. 189: 450

Family PROTODRILIDAE Czerniavsky, 1881
				Westheide, 1990: Synopses of the British Fauna # 44.
		Genus PROTODRILUS Hatschek, 1880
				von Nordheim, 1989: Zool. Scr. 18: 245-268.
			Protodrilus flabelliger Wieser, 1957

Family PROTODRILOIDAE Purschke & Jouin, 1988
				Westheide, 1990: Synopses of the British Fauna # 44.
		Genus PROTODRILOIDES
			Protodriloides chaetifer (Remane, 1926)
				Westheide, 1990: Synopses of the British Fauna 44: 112-114, f.39

Family SACCOCIRRIDAE Czerniavsky, 1881
				Westheide, 1990: Synopses of the British Fauna # 44.
		Genus SACCOCIRRUS Bobretzky, 1871
			Saccocirrus eroticus Gray, 1969

Family NERILLIDAE Levinsen, 1883
				Westheide, 1990: Synopses of the British Fauna # 44.
		Genus NERILLA O.Schmidt, 1848
			Nerilla antennata O.Schmidt, 1848
				Westheide, 1990: Synopses of the British Fauna # 44: 50-52, f.10
			Nerilla digitata Wieser, 1957
			Nerilla inopinata Gray, 1968

Family DIURODRILIDAE Kristensen & Niilonen, 1982
				Kristensen & Niilonen, 1982: Zool. Scr. 11: 1-11.
				Westheide, 1990: Synopses of the British Fauna # 44.
		Genus DIURODRILUS Remane, 1925
			Diurodrilus ankeli Ax, 1967

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